1. Nanjing University-Nanjing Forestry University Joint Institute for Plant Molecular Biology, School of Life Sciences, Nanjing University, Nanjing 210093, People’s Republic of China
2. The Key Laboratory of Western Resources Biology and Biological Technology, College of Life Sciences, Northwest University, Xi'an 710069, People’s Republic of China
3. College of Animal Science & Technology, Northwest A&F University, Yangling, Shanxi 712100, China
4. Nanjing University-Nanjing Forestry University Joint Institute for Plant Molecular Biology, Key Laboratory of Forest Genetics and Biotechnology, Nanjing Forestry University, Nanjing 210037, People’s Republic of China
5. Department of Plant and Microbial Biology, University of California, Berkeley, California 94720