1. Key Laboratory of Horticultural Plant Biology (Ministry of Education) and Key Laboratory of Horticultural Crop Biology and Genetic Improvement, Central Region (Ministry of Agriculture), Wuhan 430070, China (Y.D., Q.M., S.L., S.J., J.H., R.X., A.Z., Y.L., J.X., Q.X., Y.Z., X.D., Y.C.); and