1. Ministry of Education Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University, Lanzhou 730000, China (J.Z., X.D., T.S., J.L., S.D.R., X.G.); Department of Microbiology and Plant Biology, University of Oklahoma, Norman, Oklahoma 73019 (T.Y., X.W., S.D.R.)