1. Department of Plant Science (S.R., I.G., S.J., Y.H., D.S.L.), Intercollegiate Graduate Program in Plant Biology, Huck Institutes of Life Sciences (S.R., S.J., D.S.L.), and Department of Entomology (F.E.A., M.P., G.W.F.), Pennsylvania State University, University Park, Pennsylvania 16802;