1. Department of Botany, University of British Columbia, Vancouver, British Columbia V6T1Z4, Canada
2. Department of Botany, Graduate School of Science, Kyoto University, Oiwake-cho, Kita-Shirakawa, Sakyo-ku, Kyoto 606-8502, Japan
3. ERATO, Graduate School of Science, Nagoya University, Furo-cho, Chikusa-ku, Nagoya 464-8602, Japan
4. Department Life Sciences, Research Institute for Natural Sciences, Hanyang University, 222 Wangsipri-ro, Sungdong-gu, Seoul 133-791, Republic of Korea