1. Graduate School of Bioscience and Biotechnology, Tokyo Institute of Technology, Yokohama, Kanagawa, 226-8503, Japan
2. School of Life Science and Technology, Tokyo Institute of Technology, Yokohama, Kanagawa, 226-8503, Japan
3. RIKEN Center for Sustainable Resource Science, Yokohama, Kanagawa, 230-0045, Japan
4. Japan Science and Technology Agency, Core Research for Evolutional Science and Technology Program, Tokyo, 102-0076, Japan
5. Earth-Life Science Institute, Tokyo Institute of Technology, Tokyo, 152-8550, Japan