1. Wageningen University and Research, Bioscience, 6708 PB Wageningen, The Netherlands
2. Wageningen University and Research, Laboratory of Molecular Biology, 6708 PB Wageningen, The Netherlands
3. Enza Zaden Research and Development, 1602 DB Enkhuizen, The Netherlands
4. Max Planck Institute for Molecular Genetics, 14195 Berlin, Germany