1. Root Biology Center, Fujian Agriculture and Forestry University, Fuzhou 350002, China
2. Institute of Plant Science and Resources, Okayama University, Chuo 2-20-1, Kurashiki, Japan
3. Division of Applied Biology, Faculty of Textile Science and Technology, Shinshu University, 3-15-1, Tokida, Ueda, Nagano 386-8567, Japan
4. Crop Biotech Institute, Kyung Hee University, Yongin, Gyeonggi-do 17104, Republic of Korea