1. Division of Enzymology, Institute for Protein Research, Osaka University, 3–2 Yamadaoka, Suita, Osaka, 565–0871 Japan (Y.O., T.M., Y.K.-A., T.H.); and
2. Department of Biological Mechanisms and Functions, Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya, 464–8601 Japan (H.S., T.S.)