1. Department of Biology, Graduate School of Sciences, Kyushu University, Ropponmatsu, Fukuoka 810–8560, Japan (M.H., Z.G., H.W.);
2. Institute of Plant Biology, Biological Research Center of the Hungarian Academy of Sciences, P.O. Box 521, H–6701 Szeged, Hungary (Z.G., Z.V.);
3. Biological Laboratory, Faculty of Education, Kumamoto University, Kurokami, Kumamoto 860–0862, Japan (K.M.); and
4. School of Life Science, Tokyo University of Pharmacy and Life Science, Horinouchi, Hachioji 192–0392, Japan (N.S., M.T.)