1. Division of Plant and Crop Sciences, School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough LE12 5RD, United Kingdom (J.P.H., S.M., N.S.G., M.R.B.); Warwick HRI, University of Warwick, Wellesbourne, Warwick CV35 9EF, United Kingdom (H.C.B., R.M.H., W.P.S.); Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, United Kingdom (C.G.L., S.J.W., G.J.K.); James Hutton