1. Crop Performance and Improvement Division, Rothamsted Research, Harpenden, Herts, AL5 2JQ, United Kingdom (C.P., C.H.F.); Unilever Research, Colworth House, Sharnbrook, Bedford, MK44 1LQ, United Kingdom (J.M.F., J.E.W.); and Department of Agricultural and Environmental Science, Newcastle University, Newcastle Upon Tyne, NE1 7RU, United Kingdom (J.D.B.)