1. Zhejiang Provincial Key Laboratory of Horticultural Plant Integrative Biology, Zhejiang University, Zijingang Campus, Hangzhou 310058, People's Republic of China
2. State Agriculture Ministry Laboratory of Horticultural Plant Growth, Development, and Quality Improvement, Zhejiang University, Zijingang Campus, Hangzhou 310058, People’s Republic of China
3. State Key Laboratory of Crop Stress Biology in Arid Areas/College of Horticulture, Northwest A&F University, Yangling 712100, People's Republic of China
4. Plant and Crop Sciences Division, School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough LE125RD, United Kingdom
5. New Zealand Institute for Plant and Food Research Limited, Private Bag 92169, Auckland, New Zealand