1. Ghent University, Department of Plant Biotechnology and Bioinformatics, B-9052 Ghent, Belgium
2. VIB Center for Plant Systems Biology, B-9052 Ghent, Belgium
3. State Key Laboratory of Plant Cell and Chromosome Engineering, CAS Center for Excellence in Molecular Plant Sciences, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing 100101, China
4. University of the Chinese Academy of Sciences, Beijing 100049, China