1. Graduate School of Environmental Science, Hokkaido University, Sapporo 060-0810, Japan
2. Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo 113-8657, Japan
3. Graduate School of Agriculture, Hokkaido University, Sapporo 060-8589, Japan
4. Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Sakai 599-8531, Japan
5. Graduate School of Life Science, Hokkaido University, Sapporo 060-0810, Japan