1. School of Biological Sciences, Royal Holloway University of London, Egham, Surrey TW20 0EX, United Kingdom
2. Centre for Agricultural Research of the Hungarian Academy of Sciences, H-2462 Martonvasar, Brunszvik u. 2, Hungary
3. Institute of Biology II, BIOSS Centre for Biological Signaling Studies, and Centre for Biological Systems Analysis, University of Freiburg, 79104 Freiburg, Germany