1. Genetics, Development, and Cell Biology Department (G.K., H.Y., A.K.N., A.S.K., Y.Y., P.W.B.), Interdepartmental Genetics and Genomics Program (G.K., H.Y., A.S.K., Y.Y., P.W.B.), and Agronomy Department (P.W.B.), Iowa State University, Ames, Iowa 50011;