1. State Key Laboratory of Crop Biology, Shandong Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong 271018, China (Z.J.C., L.W., W.S., Y.Zhang, C.Z., Y.H.S., W.L., T.T.S., X.Y.Z., X.G.L., X.S.Z.); Section of Cell and Developmental Biology, University of California San Diego, La Jolla, California 92093–0116 (Y.C., Y.Zhao); and State Key Labora