1. Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo 113–8657, Japan (M.A., T.Y., Y.N., J.-i.I.); Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya 464–8601, Japan (M.N., Y.S.); RIKEN Plant Science Center, Tsurumi, Yokohama 230–0045, Japan (H.S.)