1. Department of Plant Agriculture, University of Guelph, Guelph, Canada N1G 2W1 (B.J.S.); Department of Biological Sciences, Brock University, St. Catharines, Canada L2S 3A9 (A.W.B.); and Institut des Sciences du Végétal, Centre National de la Recherche Scientifique, Gif-sur-Yvette 91 198, France (D.F.)