1. Department of Agronomy and Plant Genetics (J.L., M.F., P.M.) and Department of Plant Biology (M.G., S.I., J.S.G.), University of Minnesota, St. Paul, Minnesota 55108; United States Department of Agriculture, Agricultural Research Service, St. Paul, Minnesota 55108 (S.S.M., B.B., D.A.S., C.P.V.); and Department of Plant and Soil Sciences, Delaware Biotechnology Institute, University of Delaware, N