1. Division of Agricultural Sciences, Graduate School of Sciences and Technology for Innovation, Yamaguchi University, Yamaguchi 753–8515, Japan
2. Nagano Vegetable and Ornamental Crops Experiment Station, Shiojiri, Nagano 399–6461, Japan
3. Institute of Crop Science, National Agriculture and Food Research Organization, Tsukuba, Ibaraki 305–8518, Japan
4. Department of Biofunctional Chemistry, Graduate School of Environmental and Life Science, Okayama University, Okayama700–8530, Japan
5. Department of Agrobiological Resources, Faculty of Agriculture, Meijo University, Nagoya, Aichi 468–8502, Japan