1. National Key Laboratory of Crop Genetic Improvement, Huazhong Agricultural University, 430070 Wuhan, China
2. College of Bioengineering, Jingchu University of Technology, 448000 Jingmen, Hubei, China
3. Key Laboratory of Three Gorges Regional Plant Genetics and Germplasm Enhancement, Biotechnology Research Center, Three Gorges University, 443002 Yichang, Hubei, China
4. University Paris-Saclay, Centre National de la Recherche Scientifique, Institut National de Recherche pour L’Agriculture, L’Alimentation et L'Environnement, Institute of Plant Science of Paris-Saclay, 91405 Orsay, France