1. Department of Gene Function and Phenomics, National Institute of Genetics, Mishima, Shizuoka 411–8540, Japan
2. JST-Mirai Program, Japan Science and Technology Agency, Kawaguchi, Saitama 332–0012, Japan
3. Department of Genetics, Graduate University for Advanced Studies, SOKENDAI, Mishima, Shizuoka 411–8540, Japan