1. Nara Institute of Science and Technology, Graduate School of Biological Sciences, Ikoma, Nara 630–0192, Japan
2. University of Hamburg, Institute for Plant Sciences and Microbiology, Department of Developmental Biology, D–22609 Hamburg, Germany
3. World Premier International Research Center Initiative-Institute of Transformative Bio-Molecules (WPI-ITbM), Nagoya University, Nagoya, Aichi 464–8601, Japan
4. Institute for Advanced Research, Nagoya University, Nagoya, Aichi 464–8602, Japan