1. Marine Biological Association, The Laboratory, Citadel Hill, Plymouth PL1 2PB, United Kingdom
2. Biosciences, College of Life and Environmental Sciences, University of Exeter, Exeter, EX4 4QD United Kingdom
3. Department of Biology and Marine Biology, University of North Carolina Wilmington, Wilmington, North Carolina 28403–591
4. School of Ocean and Earth Science, University of Southampton, Southampton, SO14 3ZH, United Kingdom