1. Department of Frontier Life Sciences, Faculty of Life Sciences, Kyoto Sangyo University, Kamigamo Motoyama, Kita-ku, Kyoto 603–8555, Japan
2. Center for Plant Sciences, Kyoto Sangyo University, Kamigamo Motoyama, Kita-ku, Kyoto 603–8555, Japan
3. Department of Botany, Graduate School of Science, Kyoto University, Oiwake-cho, Kitashirakawa, Sakyo-ku, Kyoto 606–8502, Japan