1. Department of Biological Sciences, Simon Fraser University, Burnaby, British Columbia, Canada V5A 1S6 (A.B.-M., M.T., A.L.P.); Michael Smith Laboratories and Departments of Botany and Forest Sciences, University of British Columbia, Vancouver, British Columbia, Canada V6T 1Z4 (K.-A.G., D.M.M., J.B.); Canadian Forest Service, Victoria, British Columbia, Canada V8Z 1M5 (R.A.); and British Columbia