1. Department of Biological Production, Akita Prefectural University, Akita City, Akita 010–0195, Japan (N.F., M.Y., N.A., T.O., Y.N.); Core Research for Evolutional Science and Technology, Japan Science and Technology, Kawaguchi, Saitama 332–0012, Japan (N.F., M.Y., T.O., Y.N.); and National Institute of Agrobiological Sciences, Tsukuba, Ibaraki 305–8602, Japan (A.M., H.H.)