1. Center for Plant Cell Biology and Department of Botany and Plant Sciences, University of California, Riverside, California 92521
2. Department of Biology and Institute of Plant and Food Science, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China
3. Plant Genomic Network Research Team, RIKEN Center for Sustainable Resource Science, Tsurumi, Yokohama, Kanagawa 230-0045, Japan