1. Center for Plant Water-Use and Nutrition Regulation and College of Life Sciences, Joint International Research Laboratory of Water and Nutrients in Crop, Fujian Agriculture and Forestry University, Jinshan Fuzhou 350002, China
2. Institute of Oceanography, Minjiang University, Fuzhou 350108, China
3. Department of Biology, Hong Kong Baptist University and State Key Laboratory of Agrobiotechnology, The Chinese University of Hong Kong, Shatin, Hong Kong 999077, China