Thienemanniella Kieffer from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae)

Abstract

The orthoclad genus Thienemanniella Kieffer in East Asia including 30 species is reviewed. Parsimony analyses of species of Thienemanniella and Onconeura Andersen et Sæther including Corynoneurella Brundin, Tempisquitoneura Epler and Ubatubaneura Wiedenbrug et Trivinho-Strixino are performed showing that Thienemanniella afra Lehmann belongs to Corynoneurella and that other Thienemanniella species with bare eyes may belong to Onconeura. Five new species: Thienemanniella absens sp. n., T. hainanensis sp. n., T. sichuana sp. n., T. triangula sp. n., and T. wuyiensis sp. n. are described and illustrated as adult males. Thienemanniella clavicornis Kieffer, T. ginzanquerea Sasa, Kitami et Suzuki, T. nipponica (Tokunaga), T. obscura Brundin, T. togamijika Sasa et Okazawa, T. tusimufegeai Sasa et Suzuki and T. xena (Roback) are reviewed for the Chinese specimens. The holotypes of 13 Japanese species: Corynoneura ginzanquinta Sasa et Suzuki, Thienemanniella chuzeduodecima Sasa, T. ginzanquerea Sasa et Suzuki, T. nagaramaculata Sasa, T. ogasaquardecima Sasa et Suzuki, T. ogasaquindecima Sasa et Suzuki, T. okigrata Sasa, T. oyabedilata Sasa, Kawai et Ueno, T. togamijika Sasa et Okazawa, T. tonewquerea Sasa et Tanaka, T. tusimuefea Sasa et Suzuki, T. tusimufegea Sasa et Suzuki, T. yakysetea Sasa et Suzuki are re-examined. Corynoneura ginzanquinta Sasa et Suzuki is transferred to the genus Thienemanniella. A key to the males of Thienemanniella so far known is provided.