Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)

Abstract

Eleven new species of Hlavaciellus Jałoszyński, 2006 are described from Malaysia: H. adelphos, H. angustifrons, H. bifoveatus, H. cirrus, H. clandestinus, H. decoratus, H. extremalis, H. metrios, H. miser, H. peninsularis, and H. smetanai. Diagnostic characters, including head modifications of males and aedeagi are illustrated, and detailed morphology of Hlavaciellus is described for the first time on the basis of disarticulated specimens. A preliminary parsimony-based phylogenetic analysis of most currently known genera of the Cephenniini (with two genera of the Eutheiini as an outgroup) provided a robust support for the previously suggested placement of Hlavaciellus as a sister group taxon to Cephennodes Reitter, 1884. The Cephennomicrus group of genera was placed as a sister group to Cephennodes + Hlavaciellus.