1. Balthasar, V. (1937) Neue coprophage Scarabaeiden von der Ukerewe-Insel in Ost-Afrika. (5. Beitrag zur Kenntnis der Scarabaeiden der aethiopischen Region). Arbeiten ïber Morphologische und Taxonomische Entomologie aus Berlin-Dahlem, 4 (3), 192–203.
2. Boheman, C.H. (1857) Insecta Caffrariae annis 1837–1845 a J. A. Wahlberg collecta amici auxilio suffultus descripsit. Pars II. Coleoptera (Scarabaeidae). Ex officina norstedtiana, Holmiae, 395 pp.
3. Boucomont, A. (1922) Les Coléoptères Coprophages du genre Pedaria Cast. Bulletin du Muséum Nationale d’Histoire Naturelle, 28 (7), 490–495.
4. Cambefort, Y. (1991) Biogeography and evolution. In: Hanski, I. & Cambefort, Y. (Eds.), Dung beetle ecology. Princeton University Press, Princeton, New Jersey, pp. 51–68. https://doi.org/10.1515/9781400862092.51
5. Daniel, G.M & Génier, F. (2019) A preliminary checklist of the dung beetle species of Mozambique (Coleoptera: Scarabaeidae: Scarabaeinae). Catharsius La Revue, 19, 25–32.