Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s.l. (Neuroptera)

Abstract

We redescribe the holotype of Epigambria longipennis Handlirsch, 1939 from the Early Jurassic of Dobbertin (Germany), the type of the family Epigambriidae Handlirsch, 1939. Two other early Toarcian genera from Germany (Polyosmylus Ansorge, 1996 from Grimmen and Rasnitsyneura gen. nov., the type species R. aequabilis (Bode, 1953), comb. nov. from Schandelach) are assigned to Epigambriidae, which we interpret as Epigambriinae, stat. nov., a subfamily of Ithonidae s.l. The venation of Epigambriinae is most similar to that of the polystoechotid genus-group but may be distinguished from members of this group mainly by (1) a single gradate series between branches of RP in both the fore- and hind wings, and (2) the absence of crossveins between branches of MP and CuA in the forewing.