1. Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2039-2207 Main Mall, Vancouver, BC V6T 1Z4, Canada
2. Institute for the Oceans and Fisheries, University of British Columbia, 2202 Main Mall, Vancouver, BC V6T 1Z4, Canada
3. Biodiversidad y Ecología Acuática, Departamento de Zoología, Universidad de Sevilla, Avda. Reina Mercedes, 6, 41012 Sevilla, Spain
4. Hakai Institute, PO Box 309, Heriot Bay, BC V0P 1H0, Canada