1. Department of Biodiversity, Ecology and Evolution, Complutense University of Madrid (UCM), Madrid 28040, Spain
2. Department of Environmental Sciences, University of Castilla-La Mancha (UCLM), Toledo 45071, Spain
3. Department of Biodiversity and Evolutionary Biology, National Museum of Natural Sciences (MNCN), Spanish National Research Council (CSIC), Madrid 28006, Spain
4. Department of Functional Biology, University of Oviedo, Oviedo 33003, Spain