1. Institute for East China Sea Research, Organization for Marine Science and Technology, Nagasaki University, 1551-7 Taira-machi, Nagasaki 851-2213, Japan
2. Eastern Marine Biology Research Center, Fisheries Research Institute, Council of Agriculture, 22 Wuchuan Road, Taitung, Chenggong 961, Taiwan
3. Fisheries Research Institute, Council of Agriculture, 199 Hou-Ih Road, Keelung 20246, Taiwan
4. Faculty of Fisheries Sciences, Hokkaido University, 3-1-1 Minato-cho, Hakodate, Hokkaido 041-8611, Japan
5. Graduate School of Fisheries and Environmental Sciences, Nagasaki University, 1-14 Bunkyo-machi, Nagasaki 852-8521, Japan
6. Iki Fisheries Extension Center, Nagasaki Prefecture Iki Development Bureau, 1290 Ishida-cho, Iki, Nagasaki 811-5215, Japan
7. Department of Marine Biosciences, Tokyo University of Marine Science and Technology, 4-5-7 Konan, Minato, Tokyo 108-8477, Japan