1. Guangxi Laboratory on the Study of Coral Reefs in the South China Sea, Coral Reef Research Centre of China, School of Marine Sciences, Guangxi University, Nanning 530004, PR China
2. Southern Marine Science and Engineering Guangdong Laboratory, Zhuhai 519000, PR China
3. Guangxi Key Laboratory of Marine Disaster in the Beibu Gulf, Beibu Gulf University, Qinzhou 535011, PR China
4. Key Laboratory of Marine Chemistry Theory and Technology, Ministry of Education, Ocean University of China, Qingdao 266100, PR China