1. Department of Aquatic Bioscience, Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo, Tokyo 113-8657, Japan
2. International Coastal Research Center, Atmosphere and Ocean Research Institute, The University of Tokyo, 1-19-8 Otsuchi, Iwate 028-1102, Japan
3. Department of Fisheries, Faculty of Agriculture, Kindai University, 3327-204 Nakamachi, Nara 631-8505, Japan
4. Faculty of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido 060-0810, Japan
5. Institute for Marine-Earth Exploration and Engineering, Japan Agency for Marine-Earth Science and Technology, 2-15 Natsushima-cho, Yokosuka, Kanagawa 237-0061, Japan