1. Shimizu Field station, Fisheries Recourses Institute, Japan Fisheries Research and Education Agency, Shimizu, Shizuoka 424-8633, Japan
2. Environmental management unit, JAPAN NUS Co. Ltd, Shinjuku-ku, Tokyo, 160-0023 Japan
3. Nagasaki Field station, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, Nagasaki, Nagasaki 851-2213, Japan
4. Tottori Prefectural Fisheries Experimental Station, Sakai-Minato, Tottori 684-0046, Japan
5. Niigata Field station, Fisheries Resources Institute, Fisheries Research and Education Agency, Niigata, Niigata 961-8121, Japan