1. School of Biology, Faculty of Biological Sciences, University of Leeds, Leeds LS2 9JT, UK
2. Department of Biology, Dalhousie University, Halifax, Nova Scotia B3H 4R2, Canada
3. University of Guam Marine Laboratory, UOG Station, Mangilao, 96913, Guam
4. Department of Marine Resources, Weno, Chuuk State, Federated States of Micronesia
5. FSM Ridge to Reef Program, Department of Marine Resources, Weno, Chuuk State, Federated States of Micronesia
6. The Nature Conservancy X666+XG5, Kolonia, Pohnpei, Federated States of Micronesia
7. The Nature Conservancy, 4245 North Fairfax Drive, Suite 100, Arlington, Virginia 22203-1606, USA
8. Centre for Biodiversity and Conservation Science, University of Queensland, Brisbane, Queensland 4072, Australia