1. Department of Ecology and Evolution, Stony Brook University, 650 Life Sciences Building, Stony Brook, New York 11794-5245, USA
2. Department of Biology, Hobart and William Smith Colleges, 300 Pulteney St, Geneva, New York 14456, USA
3. Department of Biological Sciences, Humboldt State University, 1 Harpst St, Arcata, California 95521, USA