1. Aix Marseille Univ, Université de Toulon, CNRS, IRD, MIO UM 110, 13288 Marseille, France
2. Department of Geological Sciences, Box 112120, University of Florida, Gainesville, FL 32611-2120, USA
3. Marine and Coastal Research Laboratory, Pacific Northwest National Laboratory, 1529 W Sequim Bay Rd, Sequim, WA 98382, USA
4. School of Oceanography, University of Washington, Box 355351, Seattle, WA 98195, USA