1. Research Center of Harmful Algae and Marine Biology, and Key Laboratory of Eutrophication and Red Tide Prevention of Guangdong Higher Education Institutes, Jinan University, Guangzhou 510632, PR China
2. State Key Laboratory of Marine Environmental Science, and College of the Environment and Ecology, Xiamen University, Xiamen 361005, PR China
3. Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou 310012, PR China
4. Fourth Institute of Oceanography, Ministry of Natural Resources, Beihai 536000, PR China