1. JF Blumenbach Institute of Zoology and Anthropology, Animal Ecology, University of Göttingen, Untere Karspüle 2, 37073 Göttingen, Germany
2. Institute of Biology and Environmental Sciences, University of Oldenburg, 26129 Oldenburg, Germany
3. Institute for Chemistry and Biology of the Marine Environment (ICBM), University of Oldenburg, 26129 Oldenburg, Germany
4. Centre of Biodiversity and Sustainable Land Use, University of Göttingen, Büsgenweg 1, 37077 Göttingen, Germany