1. Natural History Museum, Cromwell Rd, Kensington, London SW7 5BD, UK
2. Departamento de Ecologia e Biologia Evolutiva, Universidade Federal de São Paulo, Rua Professor Artur Riedel, no. 275, CEP 09972-270, Diadema, SP, Brazil
3. School of Life Sciences, University of Nottingham, University Park, Nottingham NG7 2RD, UK