1. Laboratory of Quantitative Global Change Ecology, Department of Biological Sciences, University of Toronto Scarborough, Toronto, Ontario, M1C 1A4, Canada
2. 6104 Melrose Road, Cooks Creek, Manitoba, R5M 0B9, Canada
3. US Geological Survey, Alaska Science Center, Anchorage, Alaska, 99508 USA
4. Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, M5T 3B2 Canada