1. Institute of Marine Science and Technology, Shandong University, 72 Binhai Road, Qingdao 266200, PR China
2. Research Centre for Indian Ocean Ecosystem, Tianjin University of Science and Technology, Tianjin 300457, PR China
3. Tianjin Key Laboratory of Marine Resources and Chemistry, Tianjin University of Science and Technology, Tianjin 300457, PR China