1. Evolutionary Ecology Group, Department of Zoology, University of Cambridge, Cambridge CB2 3EJ, UK
2. National Marine Science Centre, Marine Ecology Research Centre, School of Environment, Science and Engineering, Southern Cross University, Coffs Harbour, New South Wales 2450, Australia
3. Fisheries NSW, NSW Department of Primary Industries, National Marine Science Centre, Coffs Harbour, New South Wales 2450, Australia
4. Fisheries NSW, NSW Department of Primary Industries, Port Stephens Fisheries Institute, Nelson Bay, New South Wales 2315, Australia